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Biographical Sketch


1973-76 B.A., Oberlin College
1977-82 Ph.D., University of North Carolina, Chapel Hill


1982-85 Postdoctoral Research Associate, University of Massachusetts, Amherst
1984 Visiting Assistant Professor, Dartmouth College
1985-88 Assistant Professor, University of California, Irvine
1988-91 Associate Professor, University of California, Irvine
1991- Hannah Professor of Microbial Ecology, Michigan State University


1988 Committee of the Public Affairs Office of the Ecological Society of America on environmental applications of genetically engineered organisms
1988-1989 National Research Council Committee on Scientific Evaluation of the Introduction of Genetically Modified Microorganisms and Plants into the Environment
1989-1997 Associate Editor, Theoretical Population Biology
1990-1992 Associate Editor, Evolution
1990-1996 National Research Council Commission on Life Sciences
1990-1996 National Research Council Board on Biology
1991-1993 Editorial Board, Microbial Ecology
1991-1999 Editorial Advisory Board, Molecular Ecology
1994 National Science Foundation Population Biology Review Panel
1995 Vice-chair, Gordon Research Conference on "Population Biology and Evolution of Microbes and their Accessory Elements"
1995-1998 Editorial Board, The American Naturalist
1996-1998 Council Member, Society for the Study of Evolution
1997 Co-chair, National Research Council Workshop on "Ecology and Evolution of Infectious Diseases"
1999 Visiting Professor for Workshop on Experimental Evolution of Bacteria, Center for Population Biology, University of California, Davis
1999 Visiting Professor for Summer Workshop on Evolutionary Biology, University of Basel, Switzerland


1976 Phi Beta Kappa
1977 National Science Foundation Graduate Fellowship
1978 Sigma Xi
1986 American Society of Naturalists President's Award for Best Paper in The American Naturalist (with B. R. Levin)
1988 National Science Foundation Presidential Young Investigator
1992 John Simon Guggenheim Memorial Foundation Fellowship
1992 American Society of Naturalists President's Award for Best Paper in The American Naturalist (with M. R. Rose, S. C. Simpson and S. C. Tadler)
1996 John D. and Catherine T. MacArthur Foundation Fellowship
1997 Elected to Fellowship in American Academy of Microbiology
1998 Elected to Fellowship in American Academy of Arts and Sciences
1999 American Society for Microbiology, Division R Lectureship (Systematics and Evolution)
2006 Elected to Membership in National Academy of Sciences.